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Dan Varland

Dan Varland


I have B.S. (1974) and M.S. (1976) degrees in Zoology from Eastern Illinois University and a Ph.D. (1991) in Animal Ecology from Iowa State University. I taught the biological sciences at community colleges in Minnesota and Iowa for 11 years after obtaining my masters. For my doctoral research, I studied the post-fledging stage in the American Kestrel. I worked from 1993 to 2009 in western Washington as the wildlife biologist for Rayonier, a large private forest landowner. My raptor work at Rayonier focused on habitat management for Northern Spotted Owls and Bald Eagles and research on Northern Goshawk nesting ecology in the managed forest. Since 1995, I have done raptor surveys and color marking on the beaches of coastal Washington. In 2009 I founded and became the Executive Director of Coastal Raptors, a non-profit organization focused on research, education and conservation programs for raptors in coastal environments.

Coastal Raptors
P.O. Box 492  
Hoquiam,  Washington  98550  United States
Phone: 360-591-5959
Email: danvarland@coastalraptors.org

   Research Interests

Research, management, and conservation of raptors in coastal and forest environments.

Species of Interest: 
  • Peregrine Falcon
  • Falco peregrinus
  • Gyrfalcon
  • Falco rusticolus
  • Merlin
  • Falco columbarius
  • American Kestrel
  • Falco sparverius
  • Bald Eagle
  • Haliaeetus leucocephalus
  • Turkey Vulture
  • Cathartes aura
  • Northern Goshawk
  • Accipiter gentilis


Varland, D.E. 2015. A capture to remember: monitoring the health of avian scavengers on the Pacific coast. Journal of Avian Medicine and Surgery 29:250-256       pdf

Varland, D.E., J.B. Buchanan, T.L. Fleming, M.K. Kenney, and T.M. Loughin. 2012. Peregrine Falcons on coastal beaches of Washington: fifteen years of banding and surveys. Journal of Raptor Research 46:57-74.        pdf

Varland, D.E., S. Ford, G. Johnson, and T. Hamer. 2012. Monitoring the health of avian scavengers on the Pacific Coast: FY2011 final draft report to the US Fish and Wildlife Service Avian Health and Disease Program, Hoquiam, WA. 57 pp.        pdf

Varland, D.E., T.L. Fleming, and J.B. Buchanan. 2008. Tundra Peregrine Falcon (Falco peregrinus tundrius) occurrence in Washington. Washington Birds 10:48-57.       pdf

Varland, D.E., L.A. Powell, M.K. Kenney, T.L. Fleming. 2008. Peregrine Falcon survival and resighting frequencies on the Washington coast, 1995-2003. Journal of Raptor Research 42:161-171.       pdf

Varland, D.E., J.A. Smallwood, L.S. Young, and M.N. Kochert. 2007. Marking techniques. Pp. 221-236 in D.M. Bird and K.L Bildstein (eds.), Raptor research and management techniques, Hancock House Publishers, Blaine, WA.       pdf

Varland, D.E. 2003. Coastal raptor surveys: 2003 final report. Unpublished report to Washington State Parks and Recreation Commission. Hoquiam, WA. 19 pp.

Finn, S.P., D.E. Varland, and J.M. Marzluff. 2002. Does Northern Goshawk breeding occupancy vary with nest-stand characteristics on the Olympic Peninsula, Washington? Journal of Raptor Research 36:265-279.       pdf

Finn, S.P., J.M. Marzluff, and D.E. Varland. 2002. Effects of landscape and local habitat attributes on Northern Goshawk site occupancy in western Washington. Forestry Science 48:427-436.       pdf

Bird, D.M., D.E. Varland, and J.J. Negro (eds.). 1996. (Book) Raptors in human landscapes. Academic Press, San Diego, CA.

Varland, D.E., E.E. Klaas, and T.M. Loughin. 1993. Use of habitat and perches, causes of mortality and time until dispersal in post-fledging American Kestrels. Journal of Field Ornithology 64:169-178.       pdf

Varland, D.E., and T.M. Loughin. 1993. Reproductive success of American Kestrels nesting along an interstate highway in central Iowa. Wilson Bulletin 105:465-474.       pdf

Varland, D.E., R.D. Andrews, and B.L. Ehresman. 1992. Establishing a nest box program for American Kestrels along an interstate highway. Iowa Department of Natural Resources Nongame Wildlife Program, Ames, IA.       pdf

Varland, D.E., and T.M. Loughin. 1992. Social hunting in broods of two and five American Kestrels after fledging. Journal of Raptor Research 26:74-80.       pdf

Varland, D.E. 1991. Behavior and ecology of post-fledging American Kestrels. Ph.D. dissertation, Iowa State University, Ames, IA.

Varland, D.E., and E.E. Klaas. 1991. Development of foraging behavior in the American Kestrel. Journal of Raptor Research 25:9-17.       pdf

Varland, D. 1982. Raptor banding and migration in the Loess Bluffs of southwest Iowa. Iowa Bird Life 52:43-47.       pdf

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